How long does it take pinyon and juniper seed to develop and how long does it persist in the soil?

In pinyon pines (Pinus monophylla and P. edulis), portions of three growing seasons are required to produce mature seeds. Pinyon seeds are short-lived with little inherent dormancy.  In Utah (Juniperus osteosperma) and western junipers (J. occidentals), it takes two growing seasons following pollination to produce mature fruits. In contrast to pinyon, juniper often have long-lived seeds (Miller et al. 2019). 

Utah juniper female cones emerging beneath the leaf scale in the spring. They will mature to full size by late summer and fall but not naturally disseminate until the following fall (photo by Rick Miller)
Utah juniper female cones in the second summer on the tree (photo by Rick Miller)
Western juniper male cones in the spring just prior to pollination. Note the exuding resin on the back of the leaf scales, which differentiates this species from Utah juniper, which lacks exuding resin ducts. (photo by Rick Miller)